| 1 (61) | Lancaster University | GBR | |
| 2 (86) | Aberystwyth University | GBR | |
| 3 (91) | University of Aberdeen | GBR | |
| 4 (107) | Natural Environment Research Council * | GBR | |
| 5 (110) | University of Leeds | GBR | |
| 6 (121) | Newcastle University, Newcastle upon Tyne | GBR | |
| 7 (123) | University of Cambridge * | GBR | |
| 8 (128) | University of Oxford | GBR | |
| 9 (143) | The University of Edinburgh * | GBR | |
| 10 (147) | The University of Manchester | GBR | |
| 11 (149) | University College London * | GBR | |
| 12 (150) | James Hutton Institute | GBR | |
| 13 (194) | University of Nottingham * | GBR | |
| 14 (200) | Bangor University | GBR | |
| 15 (201) | Biotechnology and Biological Sciences Research Council * | GBR | |
| 16 (233) | University of Exeter | GBR | |
| 17 (249) | The University of Reading | GBR | |
| 18 (259) | The University of Sheffield | GBR | |
| 19 (289) | University of Birmingham | GBR | |
| 20 (306) | Royal Botanic Gardens Kew | GBR | |
|