| 1 (33) | Cambridge University Hospitals NHS Foundation Trust | GBR |  |
| 2 (46) | University of Oxford | GBR |  |
| 3 (48) | University of Cambridge * | GBR |  |
| 4 (50) | Imperial College Healthcare NHS Trust | GBR |  |
| 5 (54) | Institute of Cancer Research | GBR |  |
| 6 (69) | Wellcome Trust Sanger Institute | GBR |  |
| 7 (80) | Cancer Research UK | GBR |  |
| 8 (83) | Imperial College London | GBR |  |
| 9 (110) | Oxford University Hospitals NHS Trust | GBR |  |
| 10 (122) | Medical Research Council | GBR |  |
| 11 (127) | European Bioinformatics Institute EMBL | GBR |  |
| 12 (138) | The University of Manchester | GBR |  |
| 13 (165) | University College London * | GBR |  |
| 14 (196) | University of Dundee | GBR |  |
| 15 (200) | University of Warwick | GBR |  |
| 16 (226) | John Innes Center | GBR |  |
| 17 (252) | London School of Hygiene and Tropical Medicine | GBR |  |
| 17 (252) | University of Bath | GBR |  |
| 19 (262) | Queen Mary, University of London | GBR |  |
| 20 (265) | University of York | GBR |  |
| 21 (280) | University of Glasgow * | GBR |  |
| 22 (281) | The University of Edinburgh * | GBR |  |
| 23 (287) | Biotechnology and Biological Sciences Research Council * | GBR |  |
| 24 (291) | AstraZeneca, United Kingdom | GBR |  |
| 25 (309) | The University of Sheffield | GBR |  |
| 26 (311) | The Francis Crick Institute | GBR |  |
| 27 (313) | Kings College London | GBR |  |
| 28 (315) | Royal Holloway - University of London | GBR |  |
| 29 (323) | University of Leeds | GBR |  |
| 30 (328) | City University London | GBR |  |
| 31 (329) | University of Southampton | GBR |  |
| 32 (335) | University of Durham | GBR |  |
| 32 (335) | Guys & Saint Thomas NHS Foundation Trust | GBR |  |
| 34 (338) | University of Bristol | GBR |  |
| 35 (343) | Loughborough University | GBR |  |
| 35 (343) | University of St. Andrews | GBR |  |
| 37 (357) | GlaxoSmithKline, United Kingdom | GBR |  |
| 38 (364) | University of Exeter | GBR |  |
| 39 (367) | University of Nottingham * | GBR |  |
| 40 (370) | University of Liverpool * | GBR |  |
| 41 (372) | Swansea University | GBR |  |
| 41 (372) | Diamond Light Source | GBR |  |
| 43 (375) | Cardiff University | GBR |  |
| 44 (376) | University of Sussex * | GBR |  |
| 45 (378) | University of Central Lancashire | GBR |  |
| 46 (380) | Manchester Academic Health Science Centre | GBR |  |
| 47 (382) | Cranfield University | GBR |  |
| 47 (382) | University of Leicester | GBR |  |
| 49 (389) | Queen's University Belfast | GBR |  |
| 49 (389) | University of Strathclyde | GBR |  |
| 51 (395) | Robert Gordon University | GBR |  |
| 52 (409) | Syngenta, United Kingdom | GBR |  |
| 52 (409) | Newcastle University, Newcastle upon Tyne | GBR |  |
| 54 (414) | University of Essex | GBR |  |
| 55 (415) | University of Birmingham | GBR |  |
| 56 (423) | University of Greenwich | GBR |  |
| 57 (438) | University of East Anglia * | GBR |  |
| 58 (439) | Lancaster University | GBR |  |
| 59 (445) | University College London Hospital NHS Trust | GBR |  |
| 60 (448) | Kingston University | GBR |  |
| 61 (458) | Northumbria University | GBR |  |
| 62 (464) | University of Huddersfield | GBR |  |
| 63 (474) | Liverpool John Moores University | GBR |  |
| 64 (479) | Natural Environment Research Council * | GBR |  |
| 65 (486) | The University of Reading | GBR |  |
| 66 (488) | Aston University | GBR |  |
| 67 (493) | Heriot-Watt University, United Kingdom | GBR |  |
| 68 (502) | Brunel University | GBR |  |
| 69 (509) | Bangor University | GBR |  |
| 70 (522) | Keele University | GBR |  |
| 71 (533) | University of Lincoln | GBR |  |
| 71 (533) | The University of Kent | GBR |  |
| 73 (535) | Science and Technology Facilities Council * | GBR |  |
| 74 (539) | The Manchester Metropolitan University | GBR |  |
| 75 (541) | Sheffield Hallam University | GBR |  |
| 76 (542) | Rutherford Appleton Laboratory | GBR |  |
| 77 (551) | University of Bradford | GBR |  |
| 78 (560) | University of Surrey | GBR |  |
| 79 (566) | De Montfort University | GBR |  |
| 79 (566) | University of Ulster | GBR |  |
| 81 (568) | Plymouth University | GBR |  |
| 82 (580) | National Physical Laboratory | GBR |  |
| 83 (592) | University of the West of England | GBR |  |
| 84 (597) | University of Aberdeen | GBR |  |
| 85 (603) | Coventry University | GBR |  |
| 86 (609) | University of Hertfordshire | GBR |  |
| 87 (613) | The Nottingham Trent University | GBR |  |
| 88 (620) | The University of Hull | GBR |  |
| 89 (646) | The Open University | GBR |  |
| 90 (682) | University of Brighton * | GBR |  |
| 91 (756) | The University of Salford | GBR |  |
|